ENVIRONMENTAL AND REGULATORY ISSUES 18.32
essential not only for the spawning success of a number of fish species, but also as habitat
for a variety of crustaceans and insects which sustain stream diversity and productivity.
Fish explore the pool and riffle sequence to locate a site suitable for spawning,
excavate a pit, deposit eggs, and, in some species, backfill with permeable gravel. After
an incubation period of 1 or 2 months, the eggs hatch and the alevins (hatchlings still
attached to their yolk sac) spend another 1 or 2 months in the intergravel environment until
the yolk sac is fully absorbed before emerging as fry. The accumulation of fine sediment
in the gravel diminishes the flow of oxygenated water and reduces the rate of removal of
metabolic wastes produced by the eggs, and dramatically reduces survival of both eggs
and alevin. Coarser sediment such as sand which fills the interstices near the surface of the
bed can trap fry within the gravel matrix, preventing their migration into open water, and
can also fill the interstices used by juvenile fish to escape from predators.
The depth of infiltration into a gravel bed by fines depends on the sizes of the fines
relative to the grains composing the bed matrix. Fine sediment is carried deep into the
bed and accumulates by plain sedimentation within the void spaces. Larger grains may
penetrate only the armor layer, creating a seal by particle bridging within the upper layer
of the subarmor. Observations in tilting flumes indicate that sedimentation within gravel
beds occurs first in the pools and the tail of bars, gradually moving upstream to the head
of the bars. The fine sediments that accumulate in gravels may be removed by large
flows which mobilize the bed and wash out the fines. This cleansing process begins at
the head of gravel bars and moves downstream, in reverse order to the clogging process
(Diplas and Parker, I985).
Gravels are maintained free of fines by high-discharge flushing flows which occur
during flood season and mobilize the bed. These large flows are also important for
maintaining the overall channel capacity and preventing the encroachment of riparian
vegetation into the active channel, which can further reduce the amount of gravel
available for spawning. Dam construction reduces or eliminates the flood flows needed to
flush finer sediment from gravel beds and maintain channel morphology, and also
eliminates the supply of gravels from upstream. These negative impacts are only partially
offset by fine sediment trapping within the reservoir, which will reduce the rate of
gravel clogging. However, in the absence of flushing flows, even low concentrations of
fine sediment will eventually clog the bed. Diminished peak flows below a dam can also
allow sediment to accumulate in pools used for rearing of fish, and will change a
complex braided stream into a single narrow channel with loss of habitat.
Flushing flows to maintain spawning gravels below dams should: (1) mobilize
gravels to flush out fines, (2) be large enough to maintain channel morphology and prevent
vegetative encroachment, and (3) prevent significant downstream transport of gravel
which would deplete spawning beds. These objectives are not complementary. Large
discharges which flush gravels and maintain the channel also transport gravels
downstream, exacerbating the problem of gravel loss and bed coarsening.
Benthic invertebrates, mainly aquatic insects having a lifespan of about 1 year, are
the major food source for anadromous fish. Because these invertebrates cannot move
from one location to another when the water level drops, the lowest flows of the year
delimit the areas where these organisms are most numerous. Continuously submerged
gravel riffles contribute the most food to fish production (Nelson et al., 1987). The lack
of flushing flow can allow encroachment by vegetation, narrowing and deepening of the
channel, affecting not only the availability of spawning gravels, but also reducing food-
producing areas and related habitat. Hydroperiod modification which desiccates the
streambed, as well as reservoir flushing that smothers this habitat, can have disastrous
consequences on benthic fauna and all species that depend on them for food.