4.4 Bedding-Plane Markings
103
as ichnofosss, or lebensspuren. Study of trace fossils constitutes the discipline of
ichnology, which has become increasingly complex since the mid-1950s and has
spawned a massive body of literature. Several of these books are listed under
"Further Reading" at the end of this chapter.
nds ace Fossils. Trace fossils are not true bodily preserved fossils; that is,
ey
do not form by conversion of a skeleton into a body fossil. They are simply
structures that originated through the activities of organisms. Interpreted broadly,
biogenic structures can be considered to clude the following: (1) bioturbation
stctures (burrows, tracks, trails, root penetration structures), (2) biostratification
suctures (algal stromatolites, graded bedding of biogenic origin), (3) bioero
sion structures (borings, scrapings, bitings), and (4) excrement (coprolites, such as
fe cal pellets or fecal castings). Not all geologists regard biostratification structures
as ace fossils, and these structures are not commonly included in published dis
ssions of trace fossils.
Trace fossils are classified into ichnogenera on the basis of characteristics
at relate to major behavioral traits of organisms and are given generic names
as Ophiomorpha. Distinctive but less important characteristics are used to
identify ichnospecies, e.g., Ophiomorpha nodosa. Trace fossils are produced by a
host of mine organisms such as crabs, flatfish, clams, molluscs, worms, shrimp,
d eel. nonmarine environments, organisms such as insects, spiders, worms,
llipedes, snails, and lizards can produce a variety of burrows and tunnels; ver
brate organisms leave tracks; and plants leave root traces. The organisms that
puce traces are rarely preserved with the traces; thus, the trace maker is com
monly
not known. Therefore, the names applied to ichnogenera and ichnospecies
geray do not refer to the trace makers themselves.
ace Fossil Assemblages. From a sedimentological standpoint, study of trace-fossil
assemblages has commonly proven to be more useful than study of individual
icogenera or ichnospecies. A trace-fossil assemblage is a basic collective term
at embraces all of the trace fossils present within a single unit of rock Although
vaous kinds of trace-fossil assemblages are recognized, groupg of trace fossils
to ichnofacies has particular significance in paleoenvironmental studies.
Seilacher (1964, 1967) introduced the concept of ichnofacies to describe associa
of trace fossils that are recurrent in time and space and that reect environ
mental conditions such as water depth (bathymetry), salinity, and the nature of
e substrate in or on which they formed (e.g., mud vs. sand bottom). Fundamen
tly, ichnofacies are sedimentary facies defined on the basis of trace fossils, and
each inofacies may include several ichnogenera.
Seilacher (1967) established six ichnofacies, which he named after character
istic inogenera. Four of these (Skolithos, Cruziana, Zoophycos, and Nereites) were
based
on e marine water depth at which they were interpreted to occur (Table
4.3; Fig. 4.38). The Glossungites ichnofacies was established for traces that occur
to hard marine surfaces, and the Scoyenia ichnofacies characterized non
marine environments. Subsequently, Frey and Seilacher (1980) established the
Tpanites ichnofacies for hardgrounds and rockgrounds; Bromley, Pemberton,
d Rahmani (1984) proposed the Te redolites ichnofacies for borgs in wood
(woodgrou
nds); and Frey and Pemberton (1987) established the Psilonichus ichno
cies for softgrounds in the marine to nonmarine environment. Several addition
icofacies have also been proposed (e.g., Bromley, 1996, p. 241); however, the
ne inofacies shown in Ta ble 4.3 are most commonly used. Sedimentologists
parcularly interested in the Skolithos, Cruziana, Nereites, and Zoophycos ichno
facies, which have the greatest potential for interpreting ancient marine environ
mtal conditions.