1.10. Matrices in Demography 27
differential or difference equations. For a simple special case, suppose two
subpopulations of sizes n
1
(t)andn
2
(t)attimet.Ifgrowthofthesub-
populations is accompanied by migration in both directions, change in the
system can be described in terms of constants a
ij
, i =1, 2,j =1, 2:
n
1
(t +1)=a
11
n
1
(t)+a
12
n
2
(t)
n
2
(t +1)=a
21
n
1
(t)+a
22
n
2
(t).
(1.10.1)
Equations 1.10.1 are identical to the matrix equation
n
1
n
2
(t +1)=
a
11
a
12
a
21
a
22
n
1
n
2
(t), (1.10.2)
which can be written compactly as
n(t +1)=An(t), (1.10.3)
where n(t) is the population vector at time t,andA is the matrix of growth
and transfer rates, referred to indifferently as a projection matrix. If the
rates a
11
, a
21
, a
12
,anda
22
are fixed over time, (1.10.3) recurrently deter-
mines the population at any arbitrary time subsequent to t. The population
at time t + k equals that at time t successively operated on k times by A:
n(t + k)=A(...(A(An(t)) ...)=A
k
n(t). (1.10.4)
If a
12
= a
21
=0,anda
11
= a
22
, the argument of Section 1.5 applies; the
two subpopulations never come into a finite, nonzero ratio to each another,
and the subgroup with the higher rate keeps growing relative to the one
with the lower rate. But if a
12
and a
21
are positive, then, no matter how
different a
11
and a
22
may be, the two subpopulations will ultimately tend
to increase at the same rate. This stability of the ratio of one population
to the other will of course occur more quickly if a
12
and a
21
are large in
relation to the difference between a
11
and a
22
.
The aspect of stability referred to above is that in which the ratio of
the sizes of the subpopulations ultimately ceases to depend on time. When
this occurs, it follows that each of the subgroups will increase at a rate
not depending on time, that is to say, in geometric progression. For when
population n
2
is c times as large as population n
1
, for all times, then, from
the first member of (1.10.1)
n
1
(t +1)=a
11
n
1
(t)+a
12
n
2
(t)=(a
11
+ ca
12
) n
1
(t),
which proves that n
1
(t) is a constant multiple of n
2
(t). This applies to any
number of subgroups, and shows for the linear model of (1.10.1) that, if
the ratio of the sizes of the subgroups to one another is constant, each
is increasing geometrically. It can then be shown that all the groups are
increasing at the same rate. Conversely, if the subgroups are increasing
geometrically and at the same rate, they are in fixed ratios to one another.
When the process of which (1.10.1) is an example attains stability, not
only are all its subpopulations increasing at the same rate and in fixed