Teredolites
ichnofacies
The Teredolites ichnofacies consists
of a characteristic assemblage of
bor-
ings in woody (xylic) substrates. These
differ from
lithic substrates in three
main ways (Bromley et al., 1984);
1)
they may be flexible instead of rigid, 2)
they are composed of combustible
material instead of mineral matter, and
3)
they are readily biodegradable.
Woodgrounds may appear in fresh-
water settings such as logjams in
fluvial cutoffs, and freshwater ichno-
coenoses consist principally of isopod
and allied borings. Because wood sub-
strates can be moved by currents, it is
important to determine whether the
borings are autochthonous (Arua,
1989) or allochthonous (Dewey and
Keady, 1987). Only the autochthonous
forms are true members of the
Teredolites
ichnofacies. These assem-
blages may also be important in
defining sequence and parasequence
boundaries.
The traces are characterized by
1)
sparse to profuse clavate (club-shaped)
borings, 2) dense excavations, but
without interpenetrating borings, 3)
walls ornamented with the texture of
the host substrate
(e.g., tree ring im-
pressions),
4)
stumpy to elongate sub-
cylindrical, subparallel excavations in
marine or marginal marine settings, 5)
shallower, sparse to profuse nonclavate
etchings (isopod borings) in freshwater
settings.
EVALUATION OF
THE
NINE RECURRING
ICHNOFACIES MODELS
These archetypal models, particularly
the marine ones, have proven to be
valuable in characterizing general envi-
ronmental conditions. In many cases,
physical sedimentary structures cannot
be used to distinguish environments
-
for example, the structures of fluvial
point bars may be very similar to those
of estuarine point bars. However, the
two settings can easily be distin-
guished because the biogenic sedi-
mentary structures are very different.
Perhaps the most misunderstood
aspect of these recurrent ichnofacies
is
their use in paleobathymetry.
Although some workers have been
complacent regarding this aspect of
environmental reconstruction (as dis-
cussed by Frey et
a/.,
1990), other ich-
nologists have long and persistently
4. TRACE FOSSILS
emphasized that local sets of environ-
mental factors are most important in
controlling the distribution of
trace-
makers, whether or not these parame-
ters occur at specific water depths. For
instance, many estuarine point bars
exhibit a high-energy, channelward
side typified by a
Skolithos association
and a low-energy
bankward side
typified by a Cruziana association
(Howard and Frey, 1985). The respec-
tive associations occur in close prox-
imity, at the same stratigraphic or
bathymetric level.
However, many environmental pa-
rameters do tend to change progres-
sively with water depth and distance
from shore
(e.g., grain size, energy lev-
els, suspended food, water turbidity),
and these gradients effect corre-
sponding changes in the distribution of
physical and biogenic
sedimentary
structures (Fig. 2). To that extent, trace
fossil associations are indeed useful in
paleobathymetry.
The long temporal duration of most
kinds of trace fossils is very impor-
tant. These basic benthic behavioral
patterns are more nearly like stable
ecologic niches than individualistic
records of particular animal species
(Frey and Seilacher, 1980, p.
202-
203). As long as the functional niche
remains advantageous under given
environmental conditions, many dif-
ferent animal species, over long inter-
vals of geological time, may be
expected to exploit it. Their preserved
traces are strikingly similar and have
equivalent significance. Hence, al-
though we conveniently and infor-
mally speak of the "Skolithos animal"
as the architect for a particular kind of
dwelling structure, numerous different
animal species actually were involved
(i.e., many biological species construct
the same type of burrow). The
longevity of recurrent ichnofacies
thereby exceeds the longevity of re-
current biofacies by a considerable
margin. Such ichnofacies are more
useful as archetypal models not only
for environmental interpretation but
also for comparisons of depositional
environments of widely differing ages.
These recurrent ichnofacies have
been designated as archetypal facies
models, with which particular local
ich-
nofacies can be compared. The arche-
types are intended to supplement, not
supplant, local ichnofacies
designa-
tions, some of which are quite distinc-
tive; corresponding ichnofaunas may
be restricted (Serna,
1986), intergrada-
tional (Marintsch and Finks, 1982), or
diverse (Dam, 1990).
The idealized ichnofacies succession
(Fig. 2) works well in most "normal" situ-
ations (Frey and Pemberton, 1984, their
fig. 5), including salinity gradients
(Bromley and Asgaard, 1991). How-
ever, nearshore assemblages can be
found in offshore sediments, and vice
versa, if the particular sediments accu-
mulated under conditions preferred by
the tracemaking organisms. The basic
controls are not physical constraints
such as water depth, distance from
shore, or some particular tectonic or
physiographic setting. Rather, they
involve substrate consistency, hydraulic
energy, rates of deposition, turbidity,
oxygen and salinity levels, toxic sub-
stances, the quality and quantity of
available food, and the ecologic or
ich-
nologic prowess of tracemakers them-
selves
(Vossler and Pemberton, 1988a).
Finally, the models should not be di-
vorced from associated patterns of
bio-
turbation. Numerous local ichnofacies,
particularly those representing
low-
energy conditions and slow rates of de-
position, are set in a totally bioturbated
texture (Fig. 13). Several generations of
burrows may be discernible via their
cross-cutting relationships, showing that
the same volume of sediment passed
repeatedly through various styles of re-
working. In environmental recon-
struction, such ichnologic fabrics may
be equally as important as the indi-
vidual, named trace fossils (Pemberton
and Frey, 1984; Howard and Frey,
1984).
PALEOENVIRONMENTAL
SIGNIFICANCE OF ICHNOFOSSILS
The application of ichnology to paleo-
environmental analysis goes far beyond
the mere establishment of general re-
curring ichnofacies. For instance,
shallow water coastal marine environ-
ments (Chapters 9, 10,
1
1,
12) consist
of many smaller sedimentological
regimes characterized by large fluctua-
tions in many physical and chemical
parameters. In order to comprehend
fully the depositional history of such
regimes, it is necessary to have reliable
means of differentiating subtle changes
in these physical and chemical parame-
ters. Physical sedimentary structures